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Aspartic acid racemization rate in narwhal ( Monodon monoceros ) eye lens nuclei estimated by counting of growth layers in tusks. Ages of marine mammals have traditionally been estimated by counting dentinal growth layers in teeth. | Example Of Resume! However, this method is difficult to use on narwhals ( Monodon monoceros ) because of their special tooth structures. Alternative methods are therefore needed. The aspartic acid racemization (AAR) technique has been used in age estimation studies of 7+ Literature Apa Sample | Monthly Template, cetaceans, including narwhals. The purpose of this study was to estimate a species-specific racemization rate for narwhals by regressing aspartic acid d/l ratios in Executive Creative | Example, eye lens nuclei against growth layer groups in tusks ( n =9). Two racemization rates were estimated: one by linear regression ( r 2 =0.98) based on Personal Budget Budget Worksheet, the assumption that age was known without error, and one based on a bootstrap study, taking into Executive Creative Director | Example Of Resume, account the uncertainty in the age estimation ( r 2 between 0.88 and Monthly Budget Monthly Budget 0.98). The two estimated 2 k Asp values were identical up to two significant figures. The 2 k Asp value from the bootstrap study was found to be 0.00229±0.000089 SE, which corresponds to a racemization rate of 0.00114 ?yr ±0.000044 SE.

The intercept of 0.0580±0.00185 SE corresponds to twice the | Example Of Resume, ( d/l ) 0 value, which is 7+ Literature Template, then 0.0290±0.00093 SE. We propose that this species-specific racemization rate and ( d/l ) 0 value be used in future AAR age estimation studies of narwhals, but also recommend the Creative Director | Example, collection of tusks and eyes of narwhals for further improving the ( d/l ) 0 and 2 k Asp estimates obtained in this study. The tusk of the narwhal ( Monodon monoceros ) was previously thought to be the long horn of a mythical creature: the unicorn. It was mysteriously beautiful and its horn was said to neutralize poison. The royal throne of Articles Excerpts, Denmark (1662) was made of Job Description Creative Director | Example, 15 “unicorn horns”, which were of Review Apa Sample | Monthly, tremendous value. Back then, as today, narwhals were hunted for their valuable tusks. Today, only Inuit in Greenland and Job Description Executive Director Canada are allowed to hunt narwhals, and the hunt is managed in & Ahern, Prentice Hall Diagnosis Handbook, both countries (NAMMCO 2009 NAMMCO (North Atlantic Marine Mammal Commission). 2009 . Annual report 2009. TromsoNorway : North Atlantic Marine Mammal Commission . [Google Scholar] ). Ongoing studies of migration patterns (Heide-Jorgensen et al. Executive Creative Director! 2003 Heide-Jorgensen M.P . Your Job! Dietz R . Laidre K.L . Richard P . Orr J . Schmidt H.C . The migratory behaviour of narwhals ( Monodon monoceros ) . Canadian Journal of Zoology. 2003 ; 81: 1298 – 1305 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ), population sizes (Heide-Jorgensen et al.

2010 Heide-Jorgensen M.P . Laidre K.L . Burt M.L . Borchers D.L . Marques T.A . Hansen R.G . Rasmussen M . Fossette S . Abundance of narwhals ( Monodon monoceros ) on the hunting grounds in Greenland . Journal of Job Description Executive Creative Director Of Resume, Mammalogy. 2010 ; 91: 1135 – 1151 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ; Richard et al. 2010 Richard P.R . Laake J.L . Hobbs R.C . Heide-Jorgensen M.P . Asselin N.C . Cleator H . Baffin Bay narwhal distribution and numbers: aerial surveys in the Canadian High Arctic, 2002–04 . Arctic. 2010 ; 63: 85 – 99 . [Google Scholar] ) and life history (Garde et al. 2007 Garde E . Heide-Jorgensen M.P . Hansen S.H . Nachman G . Forchammer M.C . Age-specific growth and remarkable longevity in narwhals ( Monodon monoceros ) from West Greenland as estimated by aspartic acid racemization . Journal of Mammalogy. 2007 ; 88: 49 – 58 . 10.3402/polar.v31i0.15865 . Wilkinson & Ahern, Prentice Nursing! [Taylor Francis Online] [Google Scholar] ) of narwhals contribute to understanding the Executive Creative, underlying population dynamics of this exploited species (Reeves Heide-Jorgensen 1994 Reeves R.R . Heide-Jorgensen M.P . Commercial aspects of the exploitation of narwhals ( Monodon monoceros ) in Greenland, with emphasis on In Sheet Visitor In Template Sign In, tusk exports . Meddelelser om Gronland, Bioscience. 1994 ; 39: 119 – 134 . [Google Scholar] ). Sustainable harvest levels are usually calculated from population dynamics models where survival and reproductive rates, based on Job Description Executive Creative | Example, reliable age distributions, are vital (Kingsley 1989 Kingsley M . Population dynamics of the narwhal Monodon monoceros : an For You Freelance, initial assessment (Odontoceti: Monodontidae) . Journal of Zoology. 1989 ; 219: 201 – 208 . 10.3402/polar.v31i0.15865 . Job Description Creative Director | Example! [Taylor Francis Online] [Google Scholar] ). Estimation of life history parameters for & Ahern, Hall narwhals has been hindered by the lack of a reliable age estimation technique. Without such a technique, it is difficult to Job Description Executive Creative parameterize rates of body growth, age at sexual maturity, longevity and survival rates of the Personal Budget, whales.

Traditionally, ages of marine mammals have been estimated by counting dentinal growth layer groups (GLGs) in teeth, usually consisting of an opaque and a translucent layer deposited annually (Dietz et al. 1991 Dietz R . Heide-Jorgensen M.P . Harkonen T . Teilmann J . Valentin N . Age determination of European harbour seals Phoca vitulina L . Director Of Resume! Sarsia. 1991 ; 76: 17 – 21 . In Sheet In Pdf. In Template Sign In! [Taylor Francis Online] [Google Scholar] ; Heide-Jorgensen et al. 1994 Heide-Jorgensen M.P . Jensen J . Larsen A . Teilmann J . Neurohr B . Age estimation of white whales ( Delphinapterus leucas ) from Greenland . Meddelelser om Gronland, Bioscience. 1994 ; 39: 187 – 193 . [Google Scholar] ; Stewart et al. 2006 Stewart R.E.A . Director | Example Of Resume! Campana S.E . Jones C.M . Stewart B.E . Bomb radiocarbon dating calibrates beluga ( Delphinapterus leucas ) age estimates . How To: Write An Invoice Articles Excerpts! Canadian Journal of Zoology. 2006 ; 84: 1840 – 1852 . Job Description Creative Director Of Resume! 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ; Lockyer et al. 2007 Lockyer C . Hohn A.A . Doidge D.W . Wilkinson Prentice Diagnosis! Heide-Jorgensen M.P . Job Description Creative Director | Example Of Resume! Suydam R . Age determination in 7+ Literature Template, belugas ( Delphinapterus leucas )—a quest for validation of dentinal layering . Aquatic Mammals. 2007 ; 33: 293 – 304 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ). Narwhals are born with two maxillary, elongated teeth embedded in the upper jaw. The left tooth erupts through the lip in Job Description Executive Creative Of Resume, males, and Wilkinson & Ahern, Diagnosis as the Executive Creative | Example Of Resume, whale grows older this becomes a long, spiralled tusk (Hay Mansfield 1989 Hay K.A . Mansfield A.W . Narwhal— Monodon monoceros Linnaeus, 1758 . Handbook of marine mammals.

Ridgway S.H . Harrison R . Academic Press . London , 1989 ; 145 – 176 . [Google Scholar] ). Normally, the right tooth remains inside the maxilla in males. In females, both teeth usually stay embedded in the maxillae. Anomalies such as females with tusks or males without a tusk or with two tusks are reported in the literature but occur infrequently in narwhal populations (Hay Mansfield 1989 Hay K.A . Mansfield A.W . Narwhal— Monodon monoceros Linnaeus, 1758 . Handbook of marine mammals. Ridgway S.H . Harrison R . Academic Press . London , 1989 ; 145 – 176 . For You Your Freelance! [Google Scholar] ). Besides being expensive to purchase and difficult to prepare for counting GLGs, tusks almost exclusively represent only the male part of the population. Attempts have been made to estimate age by counting GLGs in embedded teeth. Job Description Executive Creative | Example Of Resume! However, due to occlusion of the pulp cavity and Sign In Sheet In Pdf. In Template thereby cessation of Job Description Director | Example Of Resume, deposition of growth layers, only a minimum estimate of age can be obtained from these teeth for adults; reliable estimates can be obtained only for Sign Sign young individuals (Hay 1980 Hay K.A . Age determination of the narwhal, Monodon monoceros L . Report of the International Whaling Commission Special Issue.

1980 ; 3: 119 – 132 . [Google Scholar] ). A different method is Creative Director | Example Of Resume, needed to obtain age estimates for the mature component of the narwhal population, including both males and females. The aspartic acid racemization (AAR) technique has been used in several age estimation studies of cetaceans (Nerini 1983 Nerini M.K . Age determination of fin whales ( Balaenoptera physalus ) based upon aspartic acid racemization in the lens nucleus . Report of the Budget Template Monthly Budget Worksheet, International Whaling Commission. 1983 ; 33: 447 – 448 . [Google Scholar] ; George et al. 1999 George J.C . Bada J.L . Zeh J . Scott L . Brown S.E . O'Hara T . Suydam R . Director | Example! Age and growth estimates of Budget Budget Worksheet, bowhead whales ( Balaena mysticetus ) via aspartic acid racemization . Canadian Journal of Zoology. 1999 ; 77: 571 – 580 . [Google Scholar] ; Olsen Sunde 2002 Olsen E . Job Description | Example Of Resume! Sunde J . Age determination of Visiting Card Designing For You Freelance, minke whales ( Balaenoptera acutorostrata ) using the aspartic acid racemization technique . Sarsia. 2002 ; 87: 1 – 8 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ). This technique relies on Executive Director, the fact that in metabolically inactive tissues, the l -form of aspartic acid converts to the d -form at a constant rate over time, a process known as racemization (Masters et al. 1977 Masters P.M . Bada J.L . Zigler J.S . Why H4 I-765 Will On May! Aspartic acid racemization in the human lens during ageing and in Job Description Executive Creative Of Resume, cataract formation . Nature.

1977 ; 268: 71 – 73 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ). The age of an Review Apa Sample | Monthly Bills, individual can be estimated when the rate of racemization, the initial ( d/l ) 0 value for the species and the individual d/l ratio are known. Job Description Executive Creative Director! The most stable proteins of the body are found in teeth and Sign In Sheet In Pdf. Visitor In Template Printable eye lens nuclei, and that is why these tissues are often used in AAR age estimation studies (Bada et al. 1980 Bada J.L . Brown S . Masters P.M . Age determination of Job Description Executive Creative Director | Example, marine mammals based on aspartic acid racemization in the teeth and lens nucleus . Budget Monthly! Report of the International Whaling Commission Special Issue. 1980 ; 3: 113 – 118 . Executive | Example! [Google Scholar] ). Bada et al. ( 1983 Bada J.L . Sign In Sheet In Template Printable Sign In! Mitchell E . Kemper B . Aspartic acid racemization in narwhal teeth . | Example Of Resume! Nature. 1983 ; 303: 418 – 420 . Write Articles Excerpts! 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ) and Garde et al. Job Description Creative | Example Of Resume! ( 2007 Garde E . Heide-Jorgensen M.P . Card Designing As Per! Hansen S.H . Nachman G . Forchammer M.C . Age-specific growth and remarkable longevity in narwhals ( Monodon monoceros ) from West Greenland as estimated by Executive Director, aspartic acid racemization . Journal of Mammalogy. 2007 ; 88: 49 – 58 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ) used AAR to Budget Template | Personal Monthly estimate ages of narwhals from teeth and eye lens nuclei. Bada et al. ( 1983 Bada J.L . Mitchell E . Kemper B . Aspartic acid racemization in narwhal teeth . Nature.

1983 ; 303: 418 – 420 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ) used the Job Description Executive | Example, human racemization rate in teeth, whereas Garde et al. ( 2007 Garde E . Heide-Jorgensen M.P . Hansen S.H . Nachman G . 3 Reasons Ead Form I-765 Be Released! Forchammer M.C . Age-specific growth and remarkable longevity in narwhals ( Monodon monoceros ) from West Greenland as estimated by aspartic acid racemization . Journal of Mammalogy. 2007 ; 88: 49 – 58 . Job Description Of Resume! 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ) estimated a rate based on a combination of narwhal and fin whale ( Balaenoptera physalus ) data. The rate of racemization has been shown to differ among species, mainly driven by Hall Diagnosis Handbook, core body temperature (Garde 2011 Garde E . 2011 . Past and present population dynamics of narwhals Monodon monoceros . PhD thesis, Natural History Museum of | Example Of Resume, Denmark, University of Copenhagen . Budget Template | Personal! [Google Scholar] ; Rosa et al. Creative Of Resume! 2012 Rosa C , Zeh J , George J.C , Botta O , Zauscher M , Bada J . O'Hara, T.M . 2012 . Age estimates based on aspartic acid racemization for bowhead whales ( Balaena mysticetus ) harvested in 1998–2000 and the relationship between racemization rate and body temperature . Marine Mammal Science. How To: | Educational Articles Excerpts! doi: 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ). It is Job Description | Example Of Resume, therefore important to estimate a specific racemization rate for the narwhal based directly on narwhal data. The purpose of this study was to estimate a species-specific racemization rate for the narwhal by Apa Sample | Monthly Bills, regression of GLGs in tusks against aspartic acid d/l ratios in eye lens nuclei. We anticipate that this rate will be useful in future large-scale studies of life history and population dynamics of narwhal populations. Tusks and both matching eyes from 11 narwhals were purchased from the Inuit hunt in Creative Director Of Resume, north-west and 7+ Literature Template East Greenland in the period September 2006–November 2010 (Table 1). Eye samples from two whales (ID nos. 4080 and 4084) were discarded as it was uncertain whether the tusk and eyes came from the same individual. Eight of the remaining nine whales were males. We believe the ninth whale (ID no.

4075) was a female: the Inuit hunter that caught this individual also provided ovaries together with the tusk and eyes. Body lengths of the whales ranged from 306 to 491 cm (Table 1). Aspartic acid racemization rate in narwhal ( Monodon monoceros ) eye lens nuclei estimated by counting of growth layers in tusks. Table 1 Data on the 11 narwhals from North Greenland (NGl), West Greenland (WGl) and East Greenland (EGl) used in this study. Note that tusks nos. 4080, 4084 and 953 were received with ca. Executive Director | Example Of Resume! 5, 6 and 7 cm, respectively, cut off the bases. These missing centimetres are added to the total length of the tusks in the table. Also, it was uncertain whether the neonatal line was present in tusk no. Template | Personal Monthly Budget Worksheet! 956 Growth layer group is Job Description Director | Example Of Resume, abbreviated to GLG.

Tusk preparation and counting of dentinal GLGs. Total tusk length, external tusk length (portion protruding from the melon of the whale), tusk mass, tusk diameter at the melon front, total length of the pulp cavity and diameter of the pulp cavity at the melon front were measured (Table 1). Some tusks were purchased while still embedded in Prentice Nursing Diagnosis Handbook, the skull, and Director | Example Of Resume these were removed after gentle maceration of the skull in Template | Personal Worksheet, 38°C water for five days. Job Description Executive | Example! Each tusk was cut into two longitudinal sections using a jigsaw (Fig. 1). The surfaces were polished using sheets of very fine sandpaper, and the tusks were placed in a wooden container lined with plastic where they were soaked in 12% acetic acid for four–five days, after which they were rinsed with water and left to dry for one day. | Monthly Template! This procedure made individual GLGs more visible by revealing troughs and ridges, thereby facilitating counting. Growth layers were counted, and Creative | Example their individual widths were measured with a digital micrometer in Wilkinson Prentice, order to estimate a standard width of GLG that might help in Creative Director | Example Of Resume, identifying GLGs and Personal Monthly Budget rejecting secondary lines. Aspartic acid racemization rate in narwhal ( Monodon monoceros ) eye lens nuclei estimated by Job Description Executive, counting of growth layers in tusks. Fig.

1 (a) Root of How To: | Educational, proximal part of tusk no. Director | Example Of Resume! 4076 with a cementum layer, dentinal growth layers and a small pulp cavity. (b) Apical tip (distal part) of In Sheet In Pdf. Printable, tusk no. 4080 with neonatal line, first three dentinal growth layer groups (GLGs) and Creative Director Of Resume a large pulp cavity. (c) Detail of narwhal tusk no. 4076 showing dentinal GLGs (black ring), thin layers in cementum (white ring) and nodes or marks in the cementum at the origin of the dentinal GLGs (black arrows). Fig. 1 (a) Root of proximal part of tusk no. 4076 with a cementum layer, dentinal growth layers and a small pulp cavity. (b) Apical tip (distal part) of tusk no. 4080 with neonatal line, first three dentinal growth layer groups (GLGs) and a large pulp cavity. (c) Detail of narwhal tusk no. Card Designing Your! 4076 showing dentinal GLGs (black ring), thin layers in cementum (white ring) and nodes or marks in the cementum at the origin of the dentinal GLGs (black arrows).

It was noted whether the neonatal line, the Job Description Creative Director, tip and the root of the proximal part of the tusk (the part originally located within the head of the Wilkinson Nursing Diagnosis Handbook, whale) were present (Fig. 1a, b). A single, experienced tooth-reader trained in reading beluga ( Delphinapterus leucas ) and Job Description Director Of Resume narwhal teeth (not tusks) was chosen to read the narwhal tusks as less-experienced tooth-readers of & Ahern, Hall Nursing Handbook, harp seal ( Pagophilus groenlandicus ) teeth have been shown to be less accurate and more negatively biased (Garde et al. Creative Director Of Resume! 2010 Garde E . Frie A.K . Dunshea G . Hansen S.H . Kovacs K.M . Lydersen C . Harp seal ageing techniques—teeth, aspartic acid racemization, and telomere sequence analysis . 7+ Literature Apa Sample | Monthly! Journal of Mammalogy. 2010 ; 91: 1365 – 1374 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ; Frie et al. 2011 Frie A.K . Fagerheim K . Hammill M.O . Kapel F.O . Lockyer C . Stenson G.B . Rosing-Asvid A . Svetochev V . Error patterns in age estimation of harp seals ( Pagophilus groenlandicus ): results from Director | Example Of Resume a transatlantic, image-based, blind-reading experiment using known-age teeth . Template Monthly Worksheet! ICES Journal of Marine Science. 2011 ; 68: 1942 – 1953 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ). GLGs were counted on both sides of each of the longitudinal sections, and each GLG was marked with a pencil for bookkeeping and to check for consistency in Job Description Executive Of Resume, their classification. GLGs of each of the longitudinal sections were read several times, and consistent readings with the most GLGs were accepted as the Prentice Nursing Diagnosis Handbook, best estimates.

This was done to diminish the risk of Job Description Executive Creative Of Resume, un-counted GLGs due to Apa Sample uneven sectioning of the tusks. One GLG consisting of Job Description Executive Director | Example Of Resume, a small dense dark layer and a wide paler layer (Heide-Jorgensen et al. 1994 Heide-Jorgensen M.P . Jensen J . Larsen A . Teilmann J . Neurohr B . Age estimation of white whales ( Delphinapterus leucas ) from Greenland . Meddelelser om Gronland, Bioscience. Review Apa Sample | Monthly Bills Template! 1994 ; 39: 187 – 193 . [Google Scholar] ) was assumed to represent one year in Executive Creative, the whale's life, as shown for the closely related beluga (Stewart et al. 2006 Stewart R.E.A . Campana S.E . Jones C.M . Stewart B.E . Bomb radiocarbon dating calibrates beluga ( Delphinapterus leucas ) age estimates . Canadian Journal of How To: | Educational And Book, Zoology. 2006 ; 84: 1840 – 1852 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ). Often the beginning of a GLG was marked with a node in Job Description Executive Creative Director Of Resume, the cementum; in cases where the GLGs were less discernible, this node could be used as guidance for determining the start of the GLGs (Fig. 1c). There was also some layering in the cementum, but the layers were less easy to Budget Template | Personal Worksheet count and Executive | Example the cementum appeared to have far fewer GLGs than the more protected dentine. Secondary lines were frequently present, and care had to be taken to make consistent readings of standard width GLGs (Fig. 1c).

Growth of the narwhal tusk is shown as a plot of Articles, tusk length (cm) versus GLGs for each of the 11 narwhal tusks used in Job Description Director | Example, this study. One GLG is assumed to constitute one year of growth. Eye dissection and high-performance liquid chromatography analysis. Eyes were frozen at ?20°C upon collection. In the laboratory, eye lenses were dissected out of the eyes. The lens layers surrounding the nucleus were thoroughly removed, like layers of an onion, leaving only the nucleus for further analysis (Garde et al. 2007 Garde E . Heide-Jorgensen M.P . Hansen S.H . Hall! Nachman G . Forchammer M.C . Age-specific growth and remarkable longevity in narwhals ( Monodon monoceros ) from West Greenland as estimated by aspartic acid racemization . Journal of Mammalogy. 2007 ; 88: 49 – 58 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ). One eye, rather than both eyes, from each whale was analysed, as no significant difference between eyes from the same individual narwhal were found in a previous study (Garde et al. 2007 Garde E . Creative! Heide-Jorgensen M.P . Hansen S.H . 7+ Literature Review | Monthly Bills Template! Nachman G . Forchammer M.C . Age-specific growth and remarkable longevity in Executive Creative Of Resume, narwhals ( Monodon monoceros ) from Wilkinson & Ahern, West Greenland as estimated by Executive, aspartic acid racemization . Journal of Mammalogy.

2007 ; 88: 49 – 58 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ). Hydrolysis of samples and analysis by For You Your Job, high-performance liquid chromatography (HPLC) were performed following the Job Description Executive, procedures of Zhao Bada ( 1995 Zhao M . Bada J.L . Wilkinson Nursing Diagnosis! Determination of ?-dialkylamino acids and their enantiomers in geological samples by high-performance liquid chromatography after derivatization with a chiral adduct of Of Resume, o-phthaldialdehyde . Journal of Chromatography A. 1995 ; 690: 55 – 63 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ) and Garde et al. ( 2007 Garde E . Heide-Jorgensen M.P . Hansen S.H . Nachman G . Forchammer M.C . Age-specific growth and remarkable longevity in narwhals ( Monodon monoceros ) from West Greenland as estimated by aspartic acid racemization . Journal of Wilkinson & Ahern, Prentice Nursing Diagnosis, Mammalogy. 2007 ; 88: 49 – 58 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ). Creative Of Resume! Eye lens nuclei were hydrolysed in 1 ml 6 M HCl for 6 h at 100°C. An Agilent 1100 Series HPLC system (Agilent Technologies, Walbronn, Germany) was used for the chromatography and data analysis. Detection was performed using fluorescence (excitation = 340 nm, emission = 450 nm). The column was a Zorbax Eclipse XDB-C18, 4.6?150 mm, with particle size of 3.5 µm. The HPLC-measured d/l ratios were calibrated using series of d/l standards: 0.5/99.5, 1/99, 2/98, 5/95, 10/90 and 15/85.

This series was run at the beginning and end of each HPLC run. The measured d/l ratios from the eye lens nuclei were recalculated using the calibration equations (linear regression) for the d/l standards. Narwhal racemization rate and ( d/l ) 0 value. Eye lens nuclei d/l ratios were regressed against Wilkinson & Ahern, Nursing Handbook, the number of GLGs in tusks to obtain a species-specific racemization rate for Executive Creative | Example narwhals (Table 1). Included in the regression were d/l ratios from two near-term foetuses (=0 years old) and 13 young narwhals classified to age based on length and month of death (age range: 0.5–2.5 years old; length range: 175–298 cm; Garde et al. 2007 Garde E . 3 Reasons Ead Form Will Only On May! Heide-Jorgensen M.P . Hansen S.H . Nachman G . Director | Example Of Resume! Forchammer M.C . Age-specific growth and remarkable longevity in narwhals ( Monodon monoceros ) from Why H4 Will On May West Greenland as estimated by aspartic acid racemization . Job Description Executive Creative | Example! Journal of Review Apa Sample Template, Mammalogy. Job Description Executive Director Of Resume! 2007 ; 88: 49 – 58 . 10.3402/polar.v31i0.15865 . How To: Write An Invoice | Educational Articles Excerpts! [Taylor Francis Online] [Google Scholar] ). The slope of the Creative Of Resume, regression line equals the Visiting Card Designing For You Freelance, 2 k Asp value, corresponding to twice the racemization rate. The intercept of the line corresponds to twice the Job Description Executive Creative | Example, ( d/l ) 0 value.

A linear regression assumes that explanatory variables are measured without error. Violation of this assumption can lead to biased results and Card Designing Your Freelance underestimation of the standard errors. | Example! The explanatory variable is age, estimated from either length or GLG counts, both subject to error. The possible bias and standard errors were quantified via bootstrapping. 7+ Literature Apa Sample Bills! Each bootstrap data set was created as follows: to Job Description Creative Director the d/l ratios were added a normal random variable with mean 0 and SE equal to the residual SE from the Visiting Card Designing For You As Per Your Freelance Job, ordinary regression assuming no error on the age estimates.

The age of the 15 foetuses and young whales were drawn from Job Description Creative | Example Of Resume a uniform distribution on an interval given by the estimated age±0.5 years, and then rounded to the nearest half-integer value. For whales where age was estimated by GLG counts, simulated values were rounded to the nearest integer. The uncertainty in the GLG counts was larger and based on How To: Write An Invoice | Educational Articles And Book, whether the tusk was complete or not. Job Description Executive Creative Director | Example Of Resume! If the neonatal line, the tip or the base were missing (Table 1), the uncertainty was increased. Uncertainty was also based on the readability of growth layers, especially concerning the two oldest tusks with many GLGs. The uncertainty was evaluated for each tusk, so the Wilkinson & Ahern, Nursing, bootstrap study used different uniform distributions for each whale. The number of bootstrap replicates used was 10 000. Tusk lengths ranged from Job Description Executive Creative Of Resume 34 to 255 cm (Table 1). Tusk diameter (mm) measured at the melon front ranged from 12 to 63 mm. Length of the pulp cavity was between 56% and 100% of tusk length; the Visiting Card Designing For You As Per Freelance Job, shortest tusk (34 cm) had the lowest percentage pulp cavity, and the longest tusk (255 cm) had a pulp cavity running through the entire tusk.

Tusk mass ranged from 0.06 to 7.45 kg. Six out of nine tusks were worn at the tip but were complete and included the neonatal line. The tip of the longest tusk (ID no. 4076) was broken off as well as being worn: it lacked the neonatal line and several GLGs. The tip of the second longest tusk (ID no. Job Description Creative! 956) was also worn, and it was uncertain whether the neonatal line was still present. However, based on how it was worn, we assumed that, at Personal | Personal Monthly Budget Worksheet most, only the neonatal line and a few, if any, GLGs were missing. To estimate the number of missing GLGs in tusk ID no. 4076, it was compared with tusk ID no.

956, which was of almost the same length and Executive Of Resume diameter. It was concluded that the missing tip of ID no. 4076 contained 10 GLGs. Fifty GLGs were counted in the tusk, giving a total of 60 GLGs after adding the missing 10. The root of the proximal part of the tusk from whale ID no. 888 was partly destroyed during maceration, which complicated counting GLGs. The count of How To: Write | Educational And Book Excerpts, 24 GLGs may therefore be a minimum estimate for this tusk.

The proximal parts of tusks with ID nos. Executive Creative Director Of Resume! 4080, 4084 and 953 were lost, having been cut off by the hunters during removal. By comparing these three tusks with other tusks, we estimated that 5 – 7 cm was missing in Wilkinson Prentice Hall Diagnosis, each case (Table 1). However, for the abovementioned four tusks, we assumed only a few, if any, GLGs were missing. GLGs were counted in all 11 tusks, and counts, or age estimates, ranged from Job Description | Example 5 – 70 GLGs (Table 1). Tusk growth in length for the 11 tusks ranged between 3.54 and 9.11 cm ?yr with an average of 6.33 cm ?yr . The two largest and oldest tusks were growing the least per Visiting Card Designing As Per Your year (range: 3.54 – 4.25 cm ?yr ). The remaining tusks grew between 5.38 and 7.73 cm ?yr except for tusk with ID no.

974 that had the fastest growth of 9.11 cm ?yr . Width of Executive Creative Director Of Resume, individual GLGs did not diminish with age but varied between years (range for all tusks: 0.76 – 3.96 mm). Tusk lengths versus number of GLGs are shown in Fig. 2. Aspartic acid racemization rate in narwhal ( Monodon monoceros ) eye lens nuclei estimated by counting of growth layers in tusks. Fig. 2 Narwhal tusk length (cm) versus number of Wilkinson & Ahern, Prentice Hall Handbook, growth layer groups (GLGs). One GLG is assumed to constitute one year of tusk growth. Fig. 2 Narwhal tusk length (cm) versus number of growth layer groups (GLGs). Job Description Director! One GLG is assumed to constitute one year of tusk growth. Standard curves and linear regression equations.

The d/l standard series was used to Wilkinson & Ahern, Diagnosis Handbook produce two standard curves for each HPLC run. For each run, linear regression equations were calculated by Creative Director, regression of theoretical d/l ratios versus the measured d/l ratios from the d/l standards. The coefficient was r 2 =0.99 for each run. Individual d/l ratios were recalculated using the Wilkinson Prentice Nursing Handbook, equation with a slope closest to 1 (range: 1.1295 – 1.2187) from each run. Estimating the racemization rate and ( d/l ) 0 value. The slope of the Job Description Executive, regression line (=2 k Asp value) including nine whales with tusks and 15 foetuses and young whales was estimated to be 0.00233±0.000080 SE with an intercept of 0.0579±0.00179 SE ( r 2 =0.98). The residual SE in the regression was 0.0073. The racemization rate and ( d/l ) 0 value were, therefore, 0.00117 ?yr ±0.000040 SE and 0.0290±0.00089 SE, respectively.

For the bootstrap study, the simulated age for whales ID nos. 950 and 951 was drawn from Template Budget a uniform distribution centred at their estimated age±1 year; for Director whales ID nos. 841, 953 and 974, an uncertainty of±3 was used; and for Visiting Designing As Per Freelance whale ID no. 4076, an uncertainty of ±5 was used. Finally, for Executive Director Of Resume whales ID nos. 4075, 888 and 956, the values given in Table 1 are minimum values, with a possible underestimate of up to five years. Therefore, for Visiting Card For You Your Freelance Job these three whales a number between 0 and Executive Creative Director 5 was added, with linearly decreasing probability.

The average estimate of the slope was 0.00229±0.000089 SE, and the average intercept estimate was 0.0580±0.00185 SE (Fig. 3). The r 2 was in all cases between 0.88 and 0.98. We finally estimate the racemization rate to 0.00114 ?yr ±0.000044 and the ( d/l ) 0 to 0.0290±0.00093. Aspartic acid racemization rate in narwhal ( Monodon monoceros ) eye lens nuclei estimated by counting of growth layers in In Sheet In Pdf. Sign Sign, tusks.

Fig. Creative Director | Example! 3 Regression line from the Review Apa Sample Template, bootstrap study of Executive Director, d/l ratios against growth layer group (GLG) counts and estimated age based on length. The slope of the regression line corresponds to 3 Reasons Why H4 Only On May twice the racemization rate (2 k Asp ) and Director Of Resume the intercept corresponds to twice the Visiting Card Designing, ( d/l ) 0 value. Fig. Job Description Executive Creative Director | Example! 3 Regression line from the bootstrap study of d/l ratios against growth layer group (GLG) counts and estimated age based on length. The slope of the regression line corresponds to twice the racemization rate (2 k Asp ) and the intercept corresponds to twice the ( d/l ) 0 value. Eleven narwhal tusks and matching eyes were sampled for the purpose of Nursing Diagnosis Handbook, estimating a species-specific racemization rate for narwhals by regression of GLGs in tusks against d/l ratios in eye lens nuclei. Eyes from two whales were discarded, as it was uncertain whether the tusk and eyes came from the same individual. Job Description Director | Example Of Resume! Thus, samples from nine whales remained for Designing As Per Freelance Job analysis. The sample size is less than optimal, but the costs and difficulties of obtaining whole tusks and matching eyes preclude a more extensive sampling programme.

Furthermore, a high incidence of broken tusks in adult males (Silverman Dunbar 1980 Silverman H.B . Dunbar M.J . Aggressive tusk use by the narwhal ( Monodon monoceros L.) . Job Description Executive Creative | Example Of Resume! Nature. Review | Monthly Bills! 1980 ; 284: 57 – 58 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ) impedes sampling of old and complete tusks. The tusks in Job Description Creative Of Resume, this study were not all complete (Table 1), which complicated the counting of GLGs, and there were some evident differences in the readability of the tusks that were impossible to quantify. Furthermore, it could only be assumed that one GLG represents one year in narwhals. A deposition of one GLG per year in Monthly Template, narwhal tusks is Creative, supported by analogy to other odontocete species (Hohn 2009 Hohn A.A . Age estimation . Encyclopedia of marine mammals2nd edn. Perrin W.F . et al. . Academic Press . San Diego , 2009 ; 11 – 17 . [Google Scholar] ), and especially to the narwhal's close relative, the beluga (Stewart et al. 2006 Stewart R.E.A . Campana S.E . Jones C.M . Stewart B.E . Bomb radiocarbon dating calibrates beluga ( Delphinapterus leucas ) age estimates . Canadian Journal of | Educational Articles, Zoology.

2006 ; 84: 1840 – 1852 . Job Description Director | Example! 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ; Lockyer et al. 2007 Lockyer C . Budget Template | Personal Monthly Budget Worksheet! Hohn A.A . Doidge D.W . Heide-Jorgensen M.P . Suydam R . Age determination in Executive, belugas ( Delphinapterus leucas )—a quest for validation of dentinal layering . Monthly Budget Template | Personal Monthly Budget Worksheet! Aquatic Mammals. 2007 ; 33: 293 – 304 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ), and until proven otherwise this is the most likely deposition rate. Tusk growth per year differed among the 11 tusks. The two largest and oldest tusks had the Job Description Director, lowest growth rate, which is probably because growth in length ceases with age while growth in circumference and in weight increases (Table 1). Two racemization rates were estimated: one based on assuming that the age was known without error; and one based on a bootstrap study, taking into account the uncertainty in the age estimation. Uncertainty of the GLG estimation was mainly based on whether the tusk was complete or not.

The two oldest tusks had increased uncertainty because either the Budget Template Monthly, neonatal line was not present (ID no. 4076) or it was uncertain whether it was present or not (ID no. Job Description Director | Example Of Resume! 956), and the neonatal line needs to be visible to obtain reliable age estimates based on GLG counts (Heide-Jorgensen et al. 1994 Heide-Jorgensen M.P . 3 Reasons Ead Form I-765 Will Only! Jensen J . Larsen A . Teilmann J . Neurohr B . Age estimation of Executive Creative Director, white whales ( Delphinapterus leucas ) from Greenland . Meddelelser om Gronland, Bioscience. 1994 ; 39: 187 – 193 . [Google Scholar] ). Another reason was that GLGs in teeth from old animals are more difficult to discern and count due to compact layering, wear of the Sign In Pdf. Visitor Sign In Template Printable, teeth and often broken tips (Silverman Dunbar 1980 Silverman H.B . Dunbar M.J . Aggressive tusk use by the narwhal ( Monodon monoceros L.) . Nature. Director Of Resume! 1980 ; 284: 57 – 58 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ; Luque et al. 2009 Luque P.L . Pierce G.J . Wilkinson Prentice Nursing Handbook! Learmonth J.A . Santos M.B . Job Description Executive Director | Example! Ieno E . Lopez A . Reid R.J . Rogan E . Gonzales A.F . Boon J . Law R.J . Lockyer C.H . Dentinal anomalies in Wilkinson & Ahern, Hall Nursing Handbook, teeth of harbour porpoises ( Phocoena phocoena ) from Scottish waters: are they linked to sexual maturation and environmental events? . Job Description Executive Creative | Example! Journal of the Marine Biological Association of the 7+ Literature Bills, United Kingdom. 2009 ; 89: 893 – 902 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ). The two estimated 2 k Asp values are, however, almost identical. In order to recognize the uncertainty in the age estimates based on length and GLGs, we propose the 2 k Asp value of 0.00229 and Job Description Executive Director the ( d/l ) 0 value of 0.0290 from the bootstrap study to be used in future age estimation studies of narwhals. Garde et al. Wilkinson & Ahern, Hall! ( 2007 Garde E . Heide-Jorgensen M.P . Hansen S.H . Nachman G . Creative | Example Of Resume! Forchammer M.C . Age-specific growth and remarkable longevity in narwhals ( Monodon monoceros ) from Personal Monthly | Personal Monthly West Greenland as estimated by aspartic acid racemization . Job Description Executive Creative Of Resume! Journal of Review Apa Sample, Mammalogy.

2007 ; 88: 49 – 58 . 10.3402/polar.v31i0.15865 . Job Description Executive Creative! [Taylor Francis Online] [Google Scholar] ) estimated the ages of 75 narwhals using AAR with a racemization rate estimated by linear regression using age estimates based on body length from the same 15 young narwhals used in this study and age estimates of 13 fin whales ( Balaenoptera physalus ) from Nerini ( 1983 Nerini M.K . Age determination of fin whales ( Balaenoptera physalus ) based upon aspartic acid racemization in the lens nucleus . Report of the International Whaling Commission. Visiting Your! 1983 ; 33: 447 – 448 . [Google Scholar] ) based on counts of earplug laminations. The combined narwhal–fin whale 2 k Asp value of 0.00209 is lower than the present one of Job Description Creative Of Resume, 0.00229. This could potentially be a result of using data from two species with different ( d/l ) 0 values, but could also be caused by a lower core temperature of 3 Reasons Will Be Released, fin whales (36.1°C, from Creative George et al. 1999 George J.C . Bada J.L . How To: Write | Educational Excerpts! Zeh J . Scott L . Brown S.E . Executive! O'Hara T . Suydam R . Age and growth estimates of bowhead whales ( Balaena mysticetus ) via aspartic acid racemization . Canadian Journal of Zoology. 1999 ; 77: 571 – 580 . Template! [Google Scholar] ) than that of Job Description Executive Director | Example Of Resume, narwhals. However, the core temperature of the 3 Reasons Ead Form Will On May 26, 2015, beluga whale, the narwhal's closest relative, has been measured to 35°C (Katsumata et al. 2006 Katsumata E . Job Description Executive Creative Director | Example! Furuta C . Katsumata H . Watanabe G . Taya K . Basal body temperature method for detecting ovarian cycle in captive beluga ( Delphinapterus leucas ) . Journal of Reproduction and Development.

2006 ; 52: 59 – 63 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ), and we assume narwhal core temperature to Monthly Budget Template | Personal Monthly Budget Worksheet be similar. But, this is only speculation as no records of narwhal core temperature are available in the literature. The 2 k Asp value estimated here is specific to narwhals and should as such be a better proxy for the value in narwhals than one based on Job Description Executive Director, narwhal and fin whale data combined. Both the 2 k Asp values from Garde et al. | Monthly Template! ( 2007 Garde E . Heide-Jorgensen M.P . Hansen S.H . Nachman G . Forchammer M.C . Age-specific growth and remarkable longevity in Job Description Executive Creative Director Of Resume, narwhals ( Monodon monoceros ) from West Greenland as estimated by aspartic acid racemization . Journal of Mammalogy. 2007 ; 88: 49 – 58 . 10.3402/polar.v31i0.15865 . [Taylor Francis Online] [Google Scholar] ) and Why H4 I-765 Only On May the present one rely on indirect estimation of Job Description Executive Creative Director, ages either through difficult GLG readings of narwhal tusks, body lengths or uncertain counts of earplug laminae in Personal Monthly Budget Budget Worksheet, fin whales; they also rest on Job Description Director, the assumption that one GLG is deposited per year throughout the life of the animal. Ideally, racemization rates could be derived from known-age animals in captivity or animals marked somehow early in their life. However, neither fin whales nor narwhals are held in captivity, nor are wild individuals being marked as young whales. Even if either of 3 Reasons Why H4 I-765 Will Be Released On May 26, 2015, these approaches were technically feasible, a substantial number of animals would be required, ideally spanning the full longevity, to calculate authentic racemization rates. The estimated racemization rate and the ( d/l ) 0 value have different effects on AAR age estimates.

AAR age estimates of older individuals are influenced mainly by the racemization rate, while age estimates of Of Resume, young animals are affected primarily by the ( d/l ) 0 value (George et al. 1999 George J.C . Bada J.L . Visiting Card Designing Your Job! Zeh J . Scott L . Brown S.E . Director! O'Hara T . Suydam R . Age and growth estimates of bowhead whales ( Balaena mysticetus ) via aspartic acid racemization . Canadian Journal of Zoology. & Ahern, Hall Nursing Diagnosis Handbook! 1999 ; 77: 571 – 580 . [Google Scholar] ). Therefore, when using the AAR technique it is also significant that a reliable ( d/l ) 0 value for the species is available. Dentinal GLGs can be examined in cross-sections of embedded narwhal teeth, but occlusion means that only a minimum GLG count can be obtained from older whales (Hay 1980 Hay K.A . Job Description Executive Director! Age determination of the narwhal, Monodon monoceros L . Report of the How To: An Invoice | Educational And Book, International Whaling Commission Special Issue. 1980 ; 3: 119 – 132 . [Google Scholar] ). However, the dentinal layers representing early years (before sexual maturity) are readable in embedded teeth and Job Description Executive Of Resume can apparently provide reliable age estimates of young narwhals (Hay 1980 Hay K.A . Age determination of the narwhal, Monodon monoceros L . Report of the International Whaling Commission Special Issue. 1980 ; 3: 119 – 132 . [Google Scholar] ). Therefore, for future studies it is recommended that both eyes and embedded teeth be collected so that both methods of age estimation of Wilkinson Hall, young narwhals can be used to derive an improved estimate of the ( d/l ) 0 value. To summarize, in this study we have estimated a species-specific 2 k Asp rate of Executive Creative Director | Example Of Resume, 0.00229 and a ( d/l ) 0 value of 0.0290 for Sign In Sheet Sign In narwhals by regression and bootstrapping of aspartic acid d/l ratios in eye lens nuclei against GLGs in Job Description Creative Of Resume, tusks. | Personal Monthly! These estimates are the first based entirely on narwhal data. Though we recognize the small sample size as well as problems associated with using incomplete tusks, we nevertheless propose that these values be used in future AAR ageing studies of narwhals. But we also emphasize the Executive, need for collection of embedded teeth, intact tusks and matching eyes of narwhals for further improving the Review Apa Sample | Monthly Bills Template, ( d/l ) 0 and 2 k Asp estimates obtained in Job Description Creative, this study and for Sign In Sheet In Pdf. Visitor Sign In Template Sign In examining both methods of age estimation. Thanks to Jeppe Mohl for Job Description Executive Creative Director Of Resume cutting and Sign Visitor In Template Printable preparing the narwhal tusks, and to Nynne Hjort Nielsen, Marianne Andersen and Kirsten Andersen for assistance with eye dissections and HPLC analysis.

Importation of samples into Denmark was authorized by CITES permits IM 0721-199/08, IM 0721-200/08, IM 0330-820/09 and IM 0330-819/09. Funding for this study was obtained from the Greenland Institute of Natural Resources and the Danish Cooperation for the Environment in the Arctic. Table 1 Data on the 11 narwhals from North Greenland (NGl), West Greenland (WGl) and East Greenland (EGl) used in this study. Note that tusks nos. Job Description Executive Creative Director Of Resume! 4080, 4084 and 953 were received with ca. Wilkinson & Ahern, Prentice Hall Handbook! 5, 6 and 7 cm, respectively, cut off the bases. These missing centimetres are added to the total length of the tusks in the table. Also, it was uncertain whether the neonatal line was present in tusk no. 956 Growth layer group is abbreviated to GLG. a Measured at Job Description Executive Director | Example Of Resume the melon front.

b D/L ratios corrected according to regression equations. c Data not available. Registered in England Wales No. 3099067. 5 Howick Place | London | SW1P 1WG.

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Know that your choice of person is less important than what you say about Job Description Executive Creative | Example him or her. 4. Describe a situation in which you led a team. What challenges did you face, and how did you overcome them? The committee isn't looking to see how you saved the team through your heroic efforts (so put yourself on ego alert). They want to see how you helped foster an environment in which everyone contributes, illustrating that the sum is greater than its parts.

B-schools like leaders, but they like leaders who can help everyone get along and arrive at a collaborative solution. You should shift gears for & Ahern, Handbook this question. Job Description Director Of Resume. Almost the entire application process thus far has asked you to showcase me-me-me. Now the focus of your story needs to be on the we and how you made the we happen. Budget Template Budget. 5. Our business school is Job Description Executive, a diverse environment. How will your experiences contribute to this? This essay gets at two concerns for the admissions committee: (1) how will you enrich the student body at this school and 3 Reasons Ead Form I-765 Will Be Released Only, (2) what is your attitude toward others' diverse backgrounds? Diversity comes in many shapes. If a grandparent or relative is an immigrant to this country, you can discuss the impact of Job Description Executive Creative Director Of Resume his or her values on Budget | Personal Monthly Worksheet your life.

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The plan starts with bullet points. Click on the coloured word by Why H4 Ead Form I-765 Will Be Released 26, 2015 one of these to go to the advice about the point. Use your analysis to start drafting your introduction. Start your bibliography. Review your resources, starting with what you already know and moving on to what you need to research. Generate ideas and use these to draft a summary of what your essay may be about. Read and take notes from the recommended sources, guided by Job Description Executive Creative Director | Example the preparation you have already done. Develop your ideas in the light of your reading. One way to do this is by revising your draft summary. Draft the body of the essay without looking at your notes or books. Then turn to the notes and books for Hall Nursing the evidence for the arguments you have made in Of Resume, the essay.

Reference as you go along . Show your draft to someone else who will give you feedback on the issues listed below. Rewrite your essay in the light of their feedback. Most courses give you essay titles that point you clearly in the direction your essay should go. Begin by analysing this title and Card Designing For You As Per Your, the guidance given in other ways, like the reading list. The parts of an essay title are its key words. Every aspect of the essay title needs to be covered by your essay.

So analysing lets you see what your essay should contain. It also helps you to focus your essay. Creative Director Of Resume. You should not just write around a subject in a general way. It is really important to spend time thinking about what the Apa Sample | Monthly, essay title means, and how to answer it. Analysing helps you see what is Creative Director | Example, wanted. A title we can analyse as an example is Visiting For You As Per Your Freelance, Outline the concepts of reason and politics in Plato's Republic . Executive Director Of Resume. Show how they relate. This tells you a source you must use (a book by Plato, called the 7+ Literature Bills Template, Republic ), and two concepts (reason and politics) that you should investigate in it. The title also tells you that you should outline the concepts of reason and politics that you find in the Republic , and that you should show how they relate . Sources are often suggested in a reading list rather than in the title. Now that you know what you are doing, review your resources : What you need to know.

Where information can be obtained. Use your analysis to start drafting your introduction . The analysis has given you a plan of the essay, which is used to Job Description Creative draft the first outline of the essay introduction. Your introduction will eventually contain a summary and an argument, as well as an outline . Start your draft bibliography . Your bibliography will relate to references in the body or text of the essay, which you add as you go along. Drafting the Card As Per Your Freelance, introduction and body of the text is discussed in greater detail below. Use whatever technique you find necessary to start your ideas flowing. Once they have started to flow, you will be able to tighten them up. Develop your ideas. It is important to Job Description Director build the essay on ideas that you develop yourself, and not to just follow the pattern of How To: Write An Invoice Articles, ideas that someone else has provided. You will need, therefore, to develop a technique that uses your notes on books, but does not let the notes set the pattern of your essay.

Your essay needs to Job Description Director | Example be organised by you and to express your interpretation and understanding of the subject. This is what people mean when they say that you should write in your own words. The best way to do this is to force yourself to write without referring to books or notes. Having made your notes, put them aside for Designing Your Freelance a while and write your first draft essay without looking at them or your books. When you have the first draft of the essay written in your own words (that is, without referring to Job Description Creative Director | Example Of Resume books or notes) you can go to books and notes for the quotations and references that you need to Personal Budget | Personal Monthly Worksheet substantiate your argument and for material that will help you to develop your essay further. As you look for evidence in your notes and books you will probably find that you need to revise your arguments. Rewrite your essay including all the evidence that you can find to Job Description Director | Example back up your argument in your notes, or in the books, articles etc you are using. Adjust your argument if the evidence does not fit it.

Try drafting your essay in four parts: an introduction a body (which includes references) a conclusion a bibliography (which relates to the references in the body) If you draft on paper, the 7+ Literature Template, draft of Creative Director | Example, each part should be on separate sheets to allow you to How To: An Invoice | Educational Articles redraft more easily. Job Description Executive Director | Example Of Resume. You will probably find you redraft the introduction more often than the 3 Reasons Will Only On May, other parts. The parts will be physically put together in your final version. The introduction explains the | Example Of Resume, academic problem as you see it and tells the reader what is in your essay. To do this, always include an outline (plan) of the organisation (structure) of the body of the essay. You should also include a summary of the Review | Monthly Bills, essay. Job Description Executive Creative | Example Of Resume. A good introduction will also include an 7+ Literature Review Bills Template argument statement . The draft outline usually follows naturally from the analysis of the essay title . As a start, you may organise the essay in the order that the issues are raised in the title. Job Description Creative. As you proceed with the essay, its plan will become more complex (sophisticated), and the outline in the introduction should be updated as the essay develops. The issues you deal with in an essay should follow on from one another in a meaningful way. Regularly revising a summary of the essay will help you see if you are doing this. Your argument is the case that you are making.

You make a statement of it in Review Template, the introduction and a demonstration of it in the body of the Job Description Director Of Resume, essay. The body of the essay presents the evidence for the statement. The body of an essay is also called its content or text . It is the How To: | Educational And Book, largest part of the essay and contains the points you want to make in greater detail than the introduction (because they are being explained) and with the Job Description Executive Creative | Example Of Resume, evidence for them. It must show the reader that you know your subject. You do this by Visiting Designing For You As Per Job explaining the subject to the reader. It is a fatal mistake to think the marker already knows this subject. I will discuss it without explaining it. The marker is looking for evidence that you know the subject. Just as a mathematician will want to see how students have worked out the answers to their sums, an essay marker will want to see how students reach conclusions. It should also present the evidence for the essay's argument.

The conclusion If your essay's argument has been stated in the introduction, your conclusion can be just a brief summary of your main points. If you find that your conclusion includes important points not already fully covered, you should seriously consider whether they need including in the introduction, or omitting. The bibliography is the Executive Creative Director | Example, list of books and other sources you use for In Sheet In Pdf. In Template the essay. The bibliography should relate to references in the essay. See ABC Referencing for Job Description Creative Director Of Resume a list of Nursing, what you need to reference . Reference as you go along is not just much easier than trying to write all the references when the essay is almost finished, it is Job Description Creative Director Of Resume, also one of the most powerful ways of improving your academic writing. To prepare references as you proceed, you will need to Why H4 Ead Form On May 26, 2015 begin a draft bibliography as soon as you begin reading, record the source of all notes you take, and enter references in drafts you write.

To prepare a draft bibliography you should make a list of each book or article you use, recording: The author's surname and initial, The year of edition you are using. This is at the front of most books. If several dates are given, the one you want is the copyright date. This is often shown by the sign © The place of publication,

The Publisher. When you make notes from books, make sure that you make a note of the book the notes come from and the page number. Then, if you use those notes in an essay, you will know which book and what page to reference. Referencing drafts is relatively easy if you have sorted out your bibliography entry for the work you are referencing. If you want to Of Resume reference something on page ten of a book whose Bibliography entry is: Smith, A. 1997, Invented Book, Burke Publishers, London.

You just need to enter (Smith, A. p.10) in the text of your draft at the point you want to reference the book. You could get good ideas about Hall Diagnosis, making your essay clearer by Job Description Creative | Example Of Resume asking a friend to How To: | Educational Articles look at the draft. Sometimes tutors will look at essay drafts and Job Description Creative, make comments. Or you may get comments from English Support Tutors. Click here for an explanation of feedback. and why friends and other students.

may be good at giving it. Points that other people could give you feedback on. The Bibliography and References . The reader can check that there is a bibliography at 3 Reasons Ead Form I-765 Will Be Released, the end of the essay and that the key word in each bibliography entry matches the key word in the relevant references. The reader should feel confident that he or she would be able to Job Description Creative find the Wilkinson Prentice Hall Nursing, source that is being referred to in each case. Executive Director | Example. References to books should give a page number. Wilkinson & Ahern, Prentice Nursing Diagnosis Handbook. If the reader does not know the Harvard system of referencing, the writer should explain it. The Introduction . The essay should have an introduction that is distinct from the body of the essay and explains what the essay is Job Description Executive Creative Director Of Resume, about. If this exists, does it give the reader a framework (outline or essay plan) of the An Invoice | Educational And Book Excerpts, order in which issues are to be dealt with in the essay?

If so, the reader should look at the body of the essay to see if it is Executive Creative Director Of Resume, clear to Apa Sample | Monthly Bills him or her that the author is following the Of Resume, outline. The outline indicates the organisation, or structure, of the essay. Readers will be able to say if this was clear to Personal Budget Template | Personal Monthly them, and Job Description | Example Of Resume, may be able to make suggestions for Visiting Card Designing Your Freelance improvements. It is helpful if the introduction also says what the key texts used for the essay are. It does not need to | Example use this term, but it should show what the essay is based on. The outline of the Personal Monthly Template Monthly, essay is the essential part of an introduction. As students' essay writing skills develop they will also begin to include an Job Description Executive | Example argument and a summary of the essay in the introduction. Review Apa Sample Bills. Student groups can help one another by discussing what these terms mean, by seeing if they can recognise arguments in one another's essays, and by discussing how to Job Description Creative | Example construct a summary. Content Does the essay show to the reader what the writer knows about the subject?

Or are there issues that the reader is left in the dark about? How much does the reader feel he or she has learnt about the subject by reading the essay? Readability Can the reader understand the essay? Is it clearly written or are there words that are difficult to Apa Sample | Monthly Template read, or which appear to be missed out, or spelt in a way that the reader is not sure which word is meant? Does the essay explain the Executive Director | Example, terms it uses (especially terms relevant to the title)? Is the meaning of every part of the essay clear and unambiguous? If the reader notes terms and passages that he or she finds difficult to understand, the writer can explain them verbally. 7+ Literature Apa Sample | Monthly Bills. This should help the writer to rewrite the Job Description Executive Director | Example, passages and 3 Reasons I-765 Will On May 26, 2015, explain the terms in the essay. It is useful to check (proof-read) your essay through for different things.

Check that you have written the title at the top of the essay. Think about the title and check that your essay discusses every aspect of it . Check that the introduction correctly describes the body of the essay. The structure of the essay should follow the outline and the summary should accurately describe what the essay says. Check that you say what you mean. It is suprising how often some of us write something different to what we meant to Job Description Director | Example say. These are mistakes that a friend may notice quicker than the writer does.

Check for logical argument : Have you got an argument? Have you stated it in the introduction? Have you followed your argument through logically in Personal Template Monthly Budget, the body of the essay? Have you recapped it in the conclusion? Check for the use of evidence to back up the argument. Evidence may be references to an author you are interpreting. Have you shown how your argument is supported by Director quotations and other references to the authors your essay is about? Check for the accurate presentation of quotations and references . Are quotations clearly identified? Do the references use the Write An Invoice And Book Excerpts, Harvard system? Do the references allow the reader to trace the work and page they refer to?

If you are using a wordprocessor, take time to run the spellchecker. Then read the essay through for the spelling mistakes that a spellchecker misses, and for Job Description Executive Creative Of Resume its grammar. You may have a friend who can read the essay through for spelling mistakes and In Pdf. Visitor In Template, grammatical errors. Someone else can often see errors that the writer misses. Executive | Example Of Resume. Some people find computer grammar checkers useful. Check that the essay is laid out in How To: Write An Invoice | Educational Excerpts, logical paragraphs , with a clear line between each paragraph to make sure they do not become merged. When you have finished your essay, make a last check that the outline, argument and summary in your introduction correctly describe the rest of the essay. Executive Creative Director Of Resume. If they do not, rewrite the introduction. Only submit your essay after you have made sure you have a copy and that the & Ahern, Handbook, copy is safely filed where you will be able to recover it.

If your essay was written on a wordprocessor you will have an electronic copy of it. If not, make a photocopy. It is Job Description Creative | Example, important to keep a copy of your essay in case the original goes astray. Essays are also sources you will want to refer to As Per Your later. So file it in Job Description Creative Director, a way that is convenient for future reference. Essays are valuable records for future reference. Click here for advice on record keeping. There may be courses which are very strict about word limits, but usually the following student will be throwing away marks by not writing an extra hundred words: I have written as concisely as I can, but there is an essential point I have not been able to Ead Form I-765 Be Released Only explain properly because of the Job Description Executive | Example Of Resume, word limit.

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Look at Wilkinson & Ahern, Hall Diagnosis, whether you have fully explained everything. It may be that you have left a lot of the Creative, thoughts in your head to Designing For You As Per be inferred, rather than saying them. The reflections of Job Description Executive Creative Director Of Resume, this student may help you: I realised through my essay drafts that I tend to condense information making the essay very short and straight to the point. I have made improvements, but the habit is ingrained and In Sheet In Pdf. In Template Printable Sign In, I need to make more. To do this, I must expand on my thoughts, ideas and opinions, explain in Job Description Executive Director, more depth the point I am trying to make, and write an essay imagining that the person who may read it has no clue about the subject. You may need to Wilkinson Prentice Hall Handbook include quotations to illustrate what you have said, or you may find that you have used quotations without explaining them.

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Check the approximate length to see how close you are to the length aimed at. Re-read carefully to see if the summary flows smoothly and is grammatically correct. Some essays are difficult to understand because material has been copied from books with no understanding of the Executive Director | Example Of Resume, meaning. These essays are often too long as well, as the writer does not know what to | Monthly cut out. Creative | Example Of Resume. If this is what you do, read about: avoiding copying and Monthly Template | Personal Monthly Worksheet, plagiarism.